The total number of negative charges in the tetrapeptide, Gly-Glu-Asp-Tyr, at pH 12.5 will be . (Integer answer)

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At a highly basic pH of 12.5, all acidic protons are lost. The negative charges come from: the C-terminal carboxylate (-1), the Glu side chain carboxylate (-1), the Asp side chain carboxylate (-1), and the Tyr side chain phenoxide (-1). The N-terminal amino group is neutral (-NH2). Total = 4.

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